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Axarli et al., The N-terminal thioredoxin domain is often described as two distinct motifs: an N-terminal motif (β1α1β2) and a C-terminal motif (β3β4α3) linked by helix α2 and which together form a four β-sheet in the order 2134 with β3 anti-parallel to the others (Figure 5A)

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This was not unexpected due to the widely recognized sensitivity of bacterial movement to specific environmental conditions (Tremblay and Déziel, 2008